These scales are continually being produced by cell division in the underlying layer of the epidermis, the stratum germinativum, and the surface of individual scutes sloughs off periodically.
2.
With improved histological techniques, a second touch receptor type, Merkel cell-neurite complexes, was found in the stratum germinativum at the bottom of the epidermis, and a third, lamellated corpuscles of Vater and Pacini, was discovered in the stratum papillare of the dermis underneath the Merkel cells as published by Halata in 1975.
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